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MetaCyc Compound: Mn2+

Synonyms: manganese ion, Mn(II), Mn+2, Mn++, manganese (II) ion

Superclasses: an iona cationan inorganic cationa divalent inorganic cation
an ionan inorganic ionan inorganic cationa divalent inorganic cation

Component of:
manganese sulfate pentahydrate
manganese chloride

Chemical Formula: Mn

Molecular Weight: 54.938 Daltons

Monoisotopic Molecular Weight: 54.9380496 Daltons

SMILES: [Mn++]

InChI: InChI=1S/Mn/q+2


Unification Links: CAS:7439-96-5, CAS:16397-91-4, ChEBI:29035, ChemSpider:25916, HMDB:HMDB01333, IAF1260:33594, KEGG:C19610, PubChem:27854

Standard Gibbs Free Energy of Change Formation (ΔfG in kcal/mol): -1.2482376Inferred by computational analysis [Latendresse13]

Reactions known to consume the compound:

manganese oxidation I :
2 Mn2+ + hydrogen peroxide + 2 H+ → 2 Mn+3 + 2 H2O

manganese oxidation II :
4 Mn2+ + oxygen + 4 H+ → 4 Mn+3 + 2 H2O

In Transport reactions:
Mn2+[extracellular space] + ATP + H2O ↔ Mn2+[cytosol] + ADP + phosphate + H+,
Mn2+[periplasm] + H+[periplasm]Mn2+[cytosol] + H+[cytosol]

Enzymes activated by Mn2+, sorted by the type of activation, are:

Activator (Allosteric) of: L-fucose isomerase [Boulter73], saturated archaetidylserine synthase [Morii03], dimethylallyltranstransferase [Chen93], ω,E,E-farnesyl diphosphate synthase [Chen93], ω,E,E,E-geranyl-geranyl diphosphate synthase [Chen93], pseudaminic acid synthase [Chou05], phosphatidylserine synthase [Okada94], unsaturated archaetidylserine synthase [Morii03], phosphoglycerol geranylgeranyltransferase [Chen93a], UDP-glucose:tetrahydrobiopterin glucosyltransferase [Chung00], α-D-glucose-1-phosphate cytidylyltransferase [Thorson94], (R)-citramalyl-CoA lyase [Friedmann07], malyl-CoA lyase [Meister05], 2-nitrobenzoate nitroreductase [Iwaki07], 2-nitrophenol 2-monooxygenase [Zeyer88], inositol-1-phosphate synthase [Chen00a], glycerol-3-phosphate cytidylyltransferase [Park93], diacetyl reductase [Takusagawa01], pyruvate carboxylase [Mukhopadhyay00], carnitine monooxygenase [Ditullio94], α-acetolactate decarboxylase [Phalip94], arginase [Ikemoto89], 2,3-bisphosphoglycerate-independent phosphoglycerate mutase [vanderOost02], D-arabinose isomerase [Boulter73], N-methylhydantoin amidohydrolase [Ogawa95], nitrite-cytochrome c oxidoreductase [Tanaka83], pyruvate kinase [Schramm00], glucose dehydrogenase [Bonete96], glucokinase [Porter82], β-phosphoglucomutase [Belocopitow74], glucokinase [Hansen03], (S)-acetoin reductase [(S,S)-2,3-butanediol forming] [Takusagawa01], F420-0-glutamyl ligase [Li03], F420-1:γ-glutamyl ligase [Li03], γ-F420-2:α-L-glutamate ligase [Li03a], arginase [Schrell89], cob(I)yrinic acid a,c-diamide adenosyltransferase [Suh95], UDP-N-acetylmuramoylalanine--D-glutamate ligase [Nathenson64], 4-ureido-5-imidazolecarboxylate amidohydrolase [Rabinowitz56], pyruvate kinase [Abbe83], tartrate dehydrogenase [Kohn68], D-xylose isomerase [Kawai94], 4-hydroxy-2-oxovalerate aldolase [Powlowski93] Activator (Mechanism unknown) of: D-altronate dehydratase [Dreyer87], glutaminase B [Prusiner76], D-mannonate oxidoreductase [Portalier72], D-mannonate dehydratase [RobertBaudouy73, Dreyer87], adenosylmethionine decarboxylase [Lu07], threonine dehydrogenase [Comment 1], dipeptidyl carboxypeptidase [Chen09, Henrich93], cysteinylglycine dipeptidase [Suzuki01], cyclic di-GMP phosphodiesterase [Lacey10], proline aminopeptidase [Yoshimoto89], polyphosphate kinase [Ahn90, Haeusler92, Comment 2], GDP-glucose trehalose-6-phosphate synthase [Elbein68], GDP-D-mannose pyrophosphorylase [Shinabarger91], phosphomannose isomerase [Shinabarger91], isovitexin 7-O-xyloside 2"-O-arabinosyltransferase [Heinsbroek79], isovitexin 2"-O-arabinosyltransferase [Heinsbroek79], 5-hydroxyanthrotainin geranyltransferase [Chooi12], bacillithiol-S-transferase [Thompson14], ferrichrome synthetase [Siegmund91], 3-benzyl-3,6 -bis(cysteinylglycine)-6-(hydroxymethyl)-diketopiperazine dipeptidase [Scharf13], 1,3,6,8-tetrahydroxynaphthalene monooxygenase (quinone-forming) [Funa05], 5'-demethoxy-6-methoxypodophyllotoxin 7-O-glucosyltransferase [Berim08], 6-methoxypodophyllotoxin 7-O-glucosyltransferase [Berim08], podophyllotoxin 7-O-glucosyltransferase [Berim08], maleylpyruvate hydrolase [Liu13], cyanidin 3-O-glucoside 6"-O-rhamnosyltransferase [Kamsteeg80], delphinidin 3-O-glucoside 6"-O-rhamnosyltransferase [Kamsteeg80], 1,4-β-mannanase [Kawaguchi14], (4R)-limonene synthase [Bouwmeester98], synephrine dehydratase [Manne86], NADH kinase [Shi05a], NAD(+) kinase [Shi05a], NADH kinase [Kawai01a], NAD(+) kinase [Kawai01a], saturated archaetidylinositol phosphate synthase [Morii09], phosphoenolpyruvate phosphatase [Veljanovski06], geraniol synthase [Iijima04], hydroxyanthraquinone oxidative coupling protein [Bais03], UDPG:ginsenoside Rd glucosyltransferase [Yue05], [BtrI acyl-carrier protein]--L-glutamate ligase [Li05], 4-amino butanoyl-[BtrI acyl-carrier protein]--L-glutamate ligase [Li05], D-xylulose-5-phosphate cytidylyltransferase [Wang12b], trehalose-6-phosphate synthase [Chaudhuri09], farnesoic acid carboxyl methyltransferase [Yang06a], exotrigalacturonate lyase [Shevchik99], digalacturonate lyase [Shevchik99], peptidoglycan glycosyltransferase [Barrett05], diphosphoinositol pentakisphosphate phosphohydrolase [Leslie02], bis(diphospho)inositol tetrakisphosphate phosphohydrolase [Leslie02], adenosine tetraphosphate phosphohydrolase [Leslie02], adenosine pentaphosphate phosphohydrolase [Leslie02], diadenosine pentaphosphate phosphohydrolase [Leslie02], diadenosine hexaphosphate phosphohydrolase [Leslie02], diadenosine hexaphosphate phosphohydrolase [Leslie02], diadenosine pentaphosphate phosphohydrolase [Leslie02], adenosine pentaphosphate phosphohydrolase [Leslie02], adenosine tetraphosphate phosphohydrolase [Leslie02], diphosphoinositol pentakisphosphate phosphohydrolase [Leslie02], guanine deaminase [Yuan99], phosphatidylinositol synthase [Antonsson94], phosphatidylinositol 3-kinase [Volinia95], unsaturated archaetidylinositol phosphate synthase [Morii09], 2-phosphinomethylmalate synthase [Shimotohno88], carboxyphosphonoenolpyruvate phosphonomutase [Hidaka90], homogalacturonan methyltransferase [Goubet99], imidazoleglycerolphosphate dehydratase [Tada95], fagopyritol B1 synthase [Ueda05], fagopyritol A1 synthase [Ueda05], galactinol synthase [Ueda05], UDP-galactose:myo-inositol 3-α-D-galactosyltransferase [Ueda05], fagopyritol synthase [Ueda05], 2-keto-3-deoxy-D-glycero-D-galacto-nononate-9-phosphate synthase [Angata99], vitexin β-glucosyltransferase [Heinsbroek80], streptomycin 6-kinase [Sugiyama83], 3-hydroxymethylcephem O-carbamoyltransferase [Brewer80], histidinol dehydrogenase [Nagai91], glucuronokinase [Neufeld59], glycerol dehydrogenase [Daniel95], feruloyl-CoA:tyramine N-(hydroxycinnamoyl)transferase [Hohlfeld95], 1,3-propanediol dehydrogenase [VeigadaCunha92], 1,3-propanediol dehydrogenase [VeigadaCunha92], 1,3-propanediol dehydrogenase [Daniel95a], acetylpyruvate hydrolase [Davey75], DL-methylmalonyl-CoA racemase [Stabler85], UDP-D-glucose pyrophosphorylase [Kotake04], UDP-L-arabinose pyrophosphorylase [Kotake04], mandelate racemase [WeilMalherbe66], 1,3-propanediol dehydrogenase [Johnson87], galactokinase [Dey83], amorpha-4,11-diene synthase [Mercke00], 4-diphosphocytidyl-2C-methyl-D-erythritol synthase [Rohdich00], tyrosine 3-monooxygenase [Yamamoto01], cis-4-hydroxy-D-proline dehydrogenase [Yoneya61], D-galactonate dehydratase [Szumilo81], arabinose kinase [Neufeld60], UDP-N-acetylglucosamine pyrophosphorylase [Szumilo96], pelargonidin-3-O-glucoside-6''-O-rhamnosyltransferase [Kamsteeg80], 3-keto-5-aminohexanoate cleavage enzyme [Yorifuji77], myo-inosose-2 dehydratase [Berman66], caffeoyl-CoA:delphinidin-3,5-diglucoside 5-O-glucoside-6-O-hydroxycinnamoyltransferase [Fujiwara97], caffeoyl-CoA:delphinidin-3,5,3'-triglucoside 5-O-glucoside-6-O-hydroxycinnamoyltransferase [Fujiwara97], galactinol synthase [Liu95], 2-ketogluconokinase [NARROD56]

Enzymes inhibited by Mn2+, sorted by the type of inhibition, are:

Inhibitor (Competitive) of: L-lysine monooxygenase [Nakazawa72], glycine formimidoyltransferase [Uyeda65] Inhibitor (Mechanism unknown) of: glutaminase [Brown08], glutaminase [Brown08], phosphoglucomutase [Joshi64], methylmalonyl-CoA mutase [Bobik03], N-acetylglutamylphosphate reductase [Vogel74], imidazole glycerol phosphate synthase [Klem93], D-aminopropanol dehydrogenase [Kelley84, Campbell78], D-glucarate dehydratase [Comment 3], biotin sulfoxide reductase [Del79], diacylglycerol pyrophosphate phosphatase [Dillon96], Rnase [Goedken01], lactose 2-epimerase [Taguchi08], L-α-amino acid ligase [Kino10], L-α-amino acid ligase [Kino10], L-α-amino acid ligase [Kino09], L-α-amino acid ligase [Kino09], cellobiose 2-epimerase [Taguchi08], dihydropterin oxidase [Unnasch82], GTP cyclohydrolase [Weisberg86], glycerophosphocholine cholinephosphodiesterase [Sugimori13], UDP-glucose:cyanidin 3-O-β-D-glucosyltransferase [Ogata98], 3,7,4'-trimethylquercetagetin 6-O-methyltransferase [De85], 3,7-dimethylquercetagetin 6-O-methyltransferase [De85], 3,7,3'-methylquercetagetin 6-O-methyltransferase [De85], 7'-O-demethylcephaeline 7'-O-methyltransferase [Cheong11], omega-hydroxypalmitate O-feruloyl transferase [Rautengarten12], thiobenzoate-S-methyltransferase [Zhao12], fumigaclavine A dimethylallyltransferase [Unsold06], o-dihydric phenol O-methyltransferase (para-methylation) [Tsang79], esculetin 6-O-methyltransferase (meta-methylation) [Tsang79], phosphatidylinositol 4,5-bisphosphate lipase C [Melin92], 4-oxalocrotonate decarboxylase [Orii06], 4-oxalocrotonate tautomerase [Orii06], 2-amino-5-carboxymuconic 6-semialdehyde deaminase [Orii04], 3-phosphonopyruvate hydrolase [Kulakova03], poly(3-hydroxybutyrate) depolymerase [Handrick01], phosphatidate phosphatase [Wu96], putrescine N-methyltransferase [Hibi92], inositolphosphorylceramide synthase [Bromley03], UDP-glucose:acetone cyanohydrin β-glucosyltransferase [Hahlbrock70], homogalacturonan methyltransferase [Goubet99], 2-aminophenol 1,6-dioxygenase [Takenaka97], ε-N-trimethyllysine hydroxylase [Sachan80], indole-3-acetate carboxyl methyltransferase [Zhao07], iminodiacetate dehydrogenase [Uetz93], carboxynorspermidine decarboxylase [Nakao90], tetrahydroprotoberberine cis-N-methyltransferase [Liscombe07], desoxyhemigossypol-6-O-methyltransferase [Liu99c], deacetoxycephalosporin C hydroxylase [Baker91], deacetoxycephalosporin C hydroxylase [Dotzlaf87], deacetoxycephalosporin C synthase [Dotzlaf87], isopenicillin N synthase [Castro88], S-adenosyl-L-methionine:norcoclaurine 6-O-methyltransferase [Sato94], 2-pyrone-4,6-dicarboxylate hydrolase [Kersten82], pyruvic oxime dioxygenase [Ono96], L-lysine-α-ketoglutarate reductase [Hutzler75], phospho-β-galactosidase [Hengstenberg70], lysine N6-acetyltransferase [Bode93], L-lactate dehydrogenase [Wrba90], L-carnitine dehydrogenase [Mori88], adenylosuccinate synthetase [Ryzhova98], S-adenosyl-L-methionine: coclaurine N-methyltransferase [Choi01], choline dehydrogenase [Nagasawa76], S-adenosyl-L-methionine:isoeugenol-O-methyltransferase [Wang98a], S-adenosyl-L-methionine:eugenol-O-methyltransferase [Wang98a], UDP-glucuronate decarboxylase [Harper02], methenyl-H4MPT cyclohydrolase [DiMarco86], DIMBOA-glucoside 4-O-methyltransferase [Oikawa02], 1-sucrose:sucrose fructosyltransferase [Koops96], pectin methyltransferase [Ishikawa00], 2-pyrone-4,6-dicarboxylate hydrolase [Maruyama83], S-adenosyl-L-methionine: (S)-scoulerine-9-O-methyltransferase [Sato93], naringenin 7-O-methyltransferase [Rakwal00], Δ1-piperideine-2-carboxylate reductase [Payton82], L-sorbosone dehydrogenase [Asakura99], glutamate synthase [Schreier84], L-lysine:pyruvate aminotransferase [Schmidt87], α-D-galactosyl-(1-3)-1D-myo-inositol:raffinose galactosyltransferase [Peterbauer98], α-D-galactosyl-(1-3)-1D-myo-inositol:raffinose galactosyltransferase [Gaudreault81] Inhibitor (Other types) of: xylan 1,3-β-xylosidase [Umemoto08], β-alanine amide amidase [Komeda04], cobinamide adenosyltransferase [Buan06], geranylgeranyl diphosphate diphosphatase [Nualkaew06], raffinose synthase [Lehle73]

This compound has been characterized as a cofactor or prosthetic group of the following enzymes: 5-phospho-α-D-ribosyl 1,2-cyclic phosphate phosphodiesterase, acyl carrier protein phosphodiesterase, xylose isomerase, L-arabinose isomerase, 2-keto-3-deoxygluconokinase, glutamine synthetase, guanosine-3',5'-bis(diphosphate) 3'-diphosphatase, L-rhamnose isomerase, α-galactosidase, D-arabinose isomerase, L-fucose isomerase, α-D-ribose 1,5-phosphomutase, deoxyribose 1,5-phosphomutase, peptidoglycan glycosyltransferase, agmatinase, allantoate amidohydrolase, S-ureidoglycine aminohydrolase, UDP-N-acetylmuramate-alanine ligase, coproporphyrinogen III oxidase, L-ascorbate 6-phosphate lactonase, 2-oxopent-4-enoate hydratase, 1-deoxy-D-xylulose 5-phosphate reductoisomerase, 2C-methyl-D-erythritol 2,4-cyclodiphosphate synthase, 1-hydroxy-2-methyl-2-(E)-butenyl 4-diphosphate synthase, 3-octaprenyl-4-hydroxybenzoate carboxy-lyase, 2-succinyl-5-enolpyruvyl-6-hydroxy-3-cyclohexene-1-carboxylate synthase, 2,3-bisphosphoglycerate-independent phosphoglycerate mutase, fructose 1,6-bisphosphatase, fructose-1,6-bisphosphatase, transketolase, transketolase, isocitrate dehydrogenase, 3-isopropylmalate dehydrogenase, malate dehydrogenase, NAD-requiring, malate dehydrogenase, superoxide dismutase, histidinol dehydrogenase, histidinal dehydrogenase, phosphoribosylglycinamide formyltransferase, oligoribonuclease, RNase T exonuclease, aminopeptidase, diacylglycerol kinase, tRNA nucleotidyltransferase, 6-phospho-β-D-glucosyl-(1,4)-D-glucose glucohydrolase, monoacetylchitobiose-6-phosphate hydrolase, phenylhydantoinase, examinopeptidase, phosphoglycerol transferase, phosphoglycerol transferase, RNA ligase, glutamine synthetase deadenylase, PII-UMP deuridylylation, [protein-PII] uridylyltransferase, protein kinase, 3' to 5' proofreading exonuclease, Xaa-Pro dipeptidase, phosphodiesterase, c-di-GMP-specific, p-aminobenzoyl-glutamate hydrolase, adenine deaminase, ThDP adenylyl transferase, 8-oxo-dGTP diphosphatase, proline aminopeptidase, NADH pyrophosphatase, pyrimidine nucleotidase, dGMP 5'-phosphate phosphohydrolase, cytidine 3'-phosphate phosphohydrolase, adenosine 3'-phosphate phosphohydrolase, 3'-nucleotidase, 5'-nucleotidase, Rnase, RNase, RNA 3'-terminal phosphate cyclase, 2-deoxyglucose-6-phosphatase, ribonuclease, oxaloacetate decarboxylase, malate dehydrogenase, (S)-malate decarboxylase, dolichyl-monophosphooligosaccharide--protein glycotransferase, dolichyl-diphospho-oligosaccharide-protein glycosyltransferase, UDP-N-acetylglucosamine–dolichyl-phosphate N-acetylglucosaminyltransferase, glucuronosyl-N-acetylgalactosaminyl-proteoglycan 4-β-N-acetylgalactosaminyltransferase, glucuronosyl-N-acetylgalactosaminyl-proteoglycan 4-β-N-acetylgalactosaminyltransferase, N-acetylgalactosaminyl-proteoglycan 3-β-glucuronosyltransferase, N-acetylgalactosaminyl-proteoglycan 3-β-glucuronosyltransferase, Cys-Gly metallodipeptidase, undecaprenyldiphospho-N-acetylgalactosamine 3-β-galactosyltransferase, 7-O-acetyl-N-acetylneuraminate synthase, 3-sulfinopropionyl-CoA synthetase, succinyl-CoA synthetase (ADP-forming), acetylgalactosaminyl-O-glycosyl-glycoprotein β-1,3-N-acetylglucosaminyltransferase, 1-deoxy-D-xylulose 5-phosphate reductoisomerase, CDP-ethanolamine:ceramide ethanolamine phosphotransferase, NAD+-dependent malate dehydrogenase, oxaloacetate decarboxylase, phosphoenolpyruvate carboxykinase (GTP), pyruvate kinase, GDP-Fuc:β-D-Gal-1,3-α-D-GalNAc-1,3-α-GalNAc-diphosphoundecaprenol α-1,2-fucosyltransferase, UDP-Gal:α-D-GalNAc-1,3-α-D-GalNAc-diphosphoundecaprenol β-1,3-galactosyltransferase, 4-O-demethyl-L-noviosyl transferase, α-glucuronidase, D-psicose 3-epimerase, D-psicose 3-epimerase, D-tagatose 3-epimerase, scyllo-inosose epimerase/isomerase, nucleoside triphosphate phosphatase, thiamine diphosphate phosphatase, UTP phosphatase, ADP phosphatase, CTP diphosphatase, ATP phosphatase, 3-amino-4-hydroxybenzoic acid synthase, malic enzyme (NAD), (+)-germacrene D synthase, patchoulol synthase, β-farnesene synthase, 5-nitroanthranilate aminohydrolase, (+)-camphor 6-exo-hydroxylase, chitin synthase, lipoamidase, hydroxymethylglutaryl-CoA lyase, phosphoenolpyruvate mutase, UDP phosphatase, CTP triphosphatase, CDP-diacylglycerol:myo-inositol phosphatidyltransferase, phosphoprotein phosphatase, glucan 1,3-β-glucosidase, rhamnogalacturonan exolyase, cyclodextrin glucanotransferase, rhamnogalacturonan endolyase, β-methylmalyl-CoA lyase, (S)-malyl-CoA lyase, 2-keto-3-deoxy-L-rhamnoate aldolase, oxaloacetate acetylhydrolase, oxalate decarboxylase, oxalate oxidase, protein-S-thiocysteine sulfate hydrolase, 3-O-α-D-mannopyranosyl-α-D-mannopyranose xylosylphosphotransferase, D-erythro-3-hydroxyaspartate aldolase, D-threo-3-hydroxyaspartate aldolase, D-threo-3-hydroxyaspartate ammonia-lyase, FAD-AMP lyase (cyclizing), adenine deaminase, galactosylxylosylprotein 3-β-galactosyltransferase, UDP-galactose:β-xylose β-1,4-galactosyltransferase, UDP-D-xylose:β-D-glucoside α-1,3-D-xylosyltransferase, allantoate amidohydrolase, ureidoglycine aminohydrolase, D-arabitol 1-phosphate dehydrogenase, 3-dehydroshikimate dehydratase, 4-oxalocrotonate decarboxylase, 2-chloro-4-fluoromuconate cycloisomerase, 2-fluoromuconate cycloisomerase, 2-chloromuconate cycloisomerase, cytidine 5'-monophosphate N-acetylneuraminate synthetase, N-acetylneuraminate synthase, cytidine 5'-monophosphate N-acetylneuraminate synthetase, N-acetylneuraminate 9-phosphate synthase, 2-chloro-cis,cis-muconate cycloisomerase, kanosamine 6-kinase, dTDP-L-dihydrostreptose:streatidine-6-phosphate dihydrostreptosyltransferase, ATP:inosamime phosphotransferase, NAD-dependent ATP dehydrogenase, cis-1,4-polyisoprene:isopentenyl-diphosphate cis-1,4-isoprenetransferase, 2-succinyl-5-enolpyruvyl-6-hydroxy-3-cyclohexene-1-carboxylate synthase, O-succinylbenzoate synthase, allantoate amidohydrolase, ureidoglycolate lyase, allantoate amidohydrolase, allantoicase, (R)-linalool synthase, acetone carboxylase, gluconolactonase, 2-methyl-3-hydroxypyridine-4,5-dicarboxylate 4-decarboxylase, β-phosphoglucomutase, β-phosphoglucomutase, geraniol synthase, muconate cycloisomerase, 2-(2'-methylthio)ethylmalate synthase, cardiolipin synthase, phosphatidylglycerolphosphate synthase, phosphatidylglycerolphosphate synthase, CDP-diacylglycerol glycerol-3-phosphate 3-phosphatidyltransferase, 4-hydroxyglutamate transaminase, p-hydroxyphenylpyruvate oxidase, malate synthase, isocitrate lyase, α-ketobutyrate synthase, 2,3-dihydroxy-isovalerate dehydratase, 2,3-dihydroxy-3-methylvalerate hydro-lyase, allantoate amidohydrolase, ureidoglycolate urea-lyase, indole-3-acetyl-ala hydrolase, ureidoglycolate amidohydrolase, arginase, agmatinase, indole-3-acetyl-ala hydrolase, indole-3-acetyl-leu hydrolase, ATP-citrate lyase, enolase, acetate kinase, benzoate-CoA ligase, 4-carboxy-4-hydroxy-2-oxoadipate aldolase, 3-hydroxybenzoate-Coenzyme A ligase, hexulose-6-phosphate synthase, arginase, ATP synthase, 3-dehydroquinate synthase, formiminoglutamate formiminohydrolase, acetate kinase, creatininase, nicotinamide mononucleotide adenylyltransferase, ADP-dependent phosphofructokinase, ADP-dependent glucokinase, UDP-α-D-glucose:glycogenin α-D-glucosyltransferase, UDP-α-D-glucose:glucosyl-glycogenin α-D-glucosyltransferase, S-adenosylmethionine synthetase, L-arginine decarboxyoxidase, 2-oxoarginine decarboxylase, guanidinobutyrase, guanidinobutyrase, arginase

This compound has been characterized as an alternative cofactor or prosthetic group of the following enzymes: bis(molybdenum cofactor) guanylyltransferase, 4-diphosphocytidyl-2C-methyl-D-erythritol synthase, allantoinase, farnesyl diphosphate synthase, pyruvate kinase, NAD+ synthetase, NH3-dependent, nucleoside diphosphate kinase, NMN adenylyltransferase, methionine adenosyltransferase, folylpoly-γ-glutamate synthetase, GDP-mannose mannosyl hydrolase, isopentenyl diphosphate isomerase, ribose-phosphate diphosphokinase, glycerol kinase, phosphoribosylpyrophosphate amidotransferase, isocitrate lyase, phosphoenolpyruvate carboxykinase (ATP), cobaltochelatase, acetate kinase, oxidized nucleoside triphosphate pyrophosphohydrolase, imidazoleglycerol-phosphate dehydratase, pantothenate synthetase, adenylylsulfate kinase, molybdenum cofactor guanylyltransferase, hypoxanthine phosphoribosyltransferase, bis(molybdenum cofactor) synthase, cytosine deaminase, holo-[acyl-carrier-protein] synthase, pyruvate kinase, aspartate ammonia-lyase, carbamoyl phosphate synthetase, geranyl diphosphate synthase, dihydroneopterin triphosphate 2'-epimerase, adenine phosphoribosyltransferase, phosphonoacetate hydrolase


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